VARIABLE	DESCRIPTION	SAMPLE_TYPE	UNITS	RANGE_LOW	RANGE_HIGH	LEGAL_VALUES
ALIENNTAX	Number of non-native (alien) taxa	FISHW		0		
ALIENPIND	% alien individuals	FISHW		0	100	
ALIENPTAX	% alien taxa	FISHW		0	100	
ANOMPIND	% individuals with anomalies (% DELT anomalies)	FISHW		0	100	
BENTINVNTAX	Number of benthic invertivore taxa	FISHW				
BENTINVPIND	% benthic invertivore individuals	FISHW				
BENTINVPTAX	% benthic invertivore taxa	FISHW				
CARNNTAX	Number of carnivore taxa	FISHW				
CARNPIND	% carnivore individuals	FISHW				
CARNPTAX	% carnivore taxa	FISHW				
CATONTAX	No. Catostomid taxa	FISHW		0		
CATOPIND	% Catostomid individuals	FISHW		0	100	
CATOPTAX	% Catostomid taxa	FISHW		0	100	
CENTNTAX	Number of Centrarchidae taxa (minus Micropterus)	FISHW				
CENTPIND	% Centrarchidae individuals (minus Micropterus)	FISHW				
CENTPTAX	% Centrarchidae taxa (minus Micropterus)	FISHW				
COLDNTAX	No. coldwater taxa	FISHW		0		
COLDPIND	% coldwater individuals	FISHW		0	100	
COLDPTAX	% coldwater taxa	FISHW		0	100	
CYPRNTAX	Number of Cyprinidae taxa	FISHW				
CYPRPIND	% Cyprinidae individuals	FISHW				
CYPRPTAX	% Cyprinidae taxa	FISHW				
DATE_COL	Date of field collection					
FW_ECO3	"Three climatic regions of the US used in WSA in indicator development: Eastern Highlands, West, and Plains and Lowlands"					
FW_ECO9	Aggregated level 3 ecoregions used in WSA					
GREAT_RIVER	Great river site					
HERBNTAX	Number of herbivores taxa	FISHW	NONE	0		
HERBPIND	% herbivores individuals	FISHW	NONE	0	100	
HERBPTAX	% herbivores taxa	FISHW	NONE	0	100	
ICTANTAX	No. Ictalurid taxa	FISHW		0		
ICTAPIND	% Ictalurid individuals	FISHW		0	100	
ICTAPTAX	% Ictalurid taxa	FISHW		0	100	
INDEX_VISIT	Index visit for NRSA population estimates					
INTLINVNTAX	Number of intolerant invertivore taxa	FISHW				
INTLINVPIND	% intolerant invertivore individuals	FISHW				
INTLINVPTAX	% intolerant invertivore taxa	FISHW				
INTLLOTNTAX	Number of intolerant lotic taxa	FISHW				
INTLLOTPIND	% intolerant lotic individuals	FISHW				
INTLLOTPTAX	% intolerant lotic taxa	FISHW				
INTLMIGRNTAX	Number of intolerant migratory taxa	FISHW				
INTLMIGRPIND	% intolerant migratory individuals	FISHW				
INTLMIGRPTAX	% intolerant migratory taxa	FISHW				
INTLNTAX	No. distinct intolerant taxa based on TOLERANCE_NRSA	FISHW		0		
INTLPIND	% Intolerant individuals based on TOLERANCE_NRSA	FISHW		0	100	
INTLPTAX	% Intolerant taxa based on TOLERANCE_NRSA	FISHW		0	100	
INTLRHEONTAX	Number of intolerant rheophilic taxa	FISHW				
INTLRHEOPIND	% intolerant rheophilic individuals	FISHW				
INTLRHEOPTAX	% intolerant rheophilic taxa	FISHW				
INVNTAX	Number of invertivore taxa	FISHW				
INVPIND	% invertivore individuals	FISHW				
INVPTAX	% invertivore taxa	FISHW				
LITHNTAX	No. lithophilic taxa	FISHW		0		
LITHPIND	% lithophilic individuals	FISHW		0	100	
LITHPTAX	% lithophilic taxa	FISHW		0	100	
LOTNTAX	Number of lotic taxa	FISHW				
LOTPIND	% lotic individuals	FISHW				
LOTPTAX	% lotic taxa	FISHW				
METHOD	Method of measurement					
MIGRNTAX	Number of migratory taxa	FISHW				
MIGRPIND	% migratory individuals	FISHW				
MIGRPTAX	% migratory taxa	FISHW				
MTOLNTAX	No. distinct intermediate tolerant taxa based on TOLERANCE_NRSA	FISHW		0		
MTOLPIND	% Intermediate tolerant individuals based on TOLERANCE_NRSA	FISHW		0	100	
MTOLPTAX	% intermediate tolerant taxa based on TOLERANCE_NRSA	FISHW		0	100	
NAT_BENTINVNTAX	Number of native benthic invertivore taxa	FISHW				
NAT_BENTINVPIND	% native individuals as benthic invertivore	FISHW	NONE			
NAT_BENTINVPTAX	% native taxa as benthic invertivore	FISHW	NONE			
NAT_CARNNTAX	Number of native carnivore taxa	FISHW				
NAT_CARNPIND	% native individuals as carnivore	FISHW	NONE			
NAT_CARNPTAX	% native taxa as carnivore	FISHW	NONE			
NAT_CATONTAX	No. native Catostomid taxa	FISHW		0		
NAT_CATOPIND	% native individuals as Catostomid	FISHW	NONE	0	100	
NAT_CATOPTAX	% native taxa as Catostomid	FISHW	NONE	0	100	
NAT_CENTNTAX	Number of native Centrarchidae (minus Micropterus) taxa	FISHW				
NAT_CENTPIND	% native individuals as Centrarchidae (minus Micropterus)	FISHW	NONE			
NAT_CENTPTAX	% native taxa as Centrarchidae (minus Micropterus)	FISHW	NONE			
NAT_COLDNTAX	Number of native coldwater taxa	FISHW				
NAT_COLDPIND	% native individuals as coldwater	FISHW	NONE			
NAT_COLDPTAX	% native taxa as coldwater	FISHW	NONE			
NAT_CYPRNTAX	Number of native Cyprinidae taxa	FISHW				
NAT_CYPRPIND	% native individuals as Cyprinidae	FISHW	NONE			
NAT_CYPRPTAX	% native taxa as Cyprinidae	FISHW	NONE			
NAT_HERBNTAX	Number of native herbivores taxa	FISHW	NONE	0		
NAT_HERBPIND	% native individuals as native herbivores	FISHW	NONE	0	100	
NAT_HERBPTAX	% native taxa as native herbivores	FISHW	NONE	0	100	
NAT_ICTANTAX	Number of native Ictaluridae taxa	FISHW				
NAT_ICTAPIND	% native individuals as Ictaluridae	FISHW	NONE			
NAT_ICTAPTAX	% native taxa as Ictaluridae	FISHW	NONE			
NAT_INTLINVNTAX	Number of native intolerant invertivore taxa	FISHW				
NAT_INTLINVPIND	% native individuals as intolerant invertivore	FISHW	NONE			
NAT_INTLINVPTAX	% native taxa as intolerant invertivore	FISHW	NONE			
NAT_INTLLOTNTAX	Number of native intolerant lotic taxa	FISHW				
NAT_INTLLOTPIND	% native individuals as intolerant lotic	FISHW	NONE			
NAT_INTLLOTPTAX	% native taxa as intolerant lotic	FISHW	NONE			
NAT_INTLMIGRNTAX	Number of native intolerant migratory taxa	FISHW				
NAT_INTLMIGRPIND	% native individuals as intolerant migratory	FISHW	NONE			
NAT_INTLMIGRPTAX	% native taxa as intolerant migratory	FISHW	NONE			
NAT_INTLNTAX	No. distinct intolerant native taxa based on TOLERANCE_NRSA	FISHW		0		
NAT_INTLPIND	% Intolerant native individuals based on TOLERANCE_NRSA	FISHW		0	100	
NAT_INTLPTAX	% Intolerant native taxa based on TOLERANCE_NRSA	FISHW		0	100	
NAT_INTLRHEONTAX	Number of native intolerant rheophilic taxa	FISHW				
NAT_INTLRHEOPIND	% native individuals as intolerant rheophilic	FISHW	NONE			
NAT_INTLRHEOPTAX	% native taxa as intolerant rheophilic	FISHW	NONE			
NAT_INVNTAX	Number of native invertivores taxa	FISHW				
NAT_INVPIND	% native individuals as invertivores	FISHW	NONE			
NAT_INVPTAX	% native taxa as invertivores	FISHW	NONE			
NAT_LITHNTAX	Number of native simple lithophils taxa	FISHW				
NAT_LITHPIND	% native individuals as simple lithophils	FISHW	NONE			
NAT_LITHPTAX	% native taxa as simple lithophils	FISHW	NONE			
NAT_LOTNTAX	Number of native lotic taxa	FISHW				
NAT_LOTPIND	% native individuals as lotic	FISHW	NONE			
NAT_LOTPTAX	% native taxa as lotic	FISHW	NONE			
NAT_MIGRNTAX	No. native migratory taxa	FISHW		0		
NAT_MIGRPIND	% native individuals as migratory	FISHW	NONE	0	100	
NAT_MIGRPTAX	% native taxa as migratory	FISHW	NONE	0	100	
NAT_MTOLNTAX	No. distinct intermediate tolerant native taxa based on TOLERANCE_NRSA	FISHW		0		
NAT_MTOLPIND	% Intermediate tolerant native individuals based on TOLERANCE_NRSA	FISHW		0	100	
NAT_MTOLPTAX	% intermediate tolerant native taxa based on TOLERANCE_NRSA	FISHW		0	100	
NAT_NTOLBENTNTAX	No. native non-tolerant benthic taxa based on TOLERANCE_NRSA	FISHW		0		
NAT_NTOLBENTPIND	% native non-tolerant benthic individuals based on TOLERANCE_NRSA	FISHW		0	100	
NAT_NTOLBENTPTAX	% native non-tolerant benthic taxa based on TOLERANCE_NRSA	FISHW		0	100	
NAT_NTOLCARNNTAX	No. native non-tolerant carnivore taxa based on TOLERANCE_NRSA	FISHW				
NAT_NTOLCARNPIND	% native non-tolerant carnivore individuals based on TOLERANCE_NRSA	FISHW				
NAT_NTOLCARNPTAX	% native non-tolerant carnivore taxa based on TOLERANCE_NRSA	FISHW				
NAT_NTOLINVNTAX	No. native non-tolerant invertivore taxa based on TOLERANCE_NRSA	FISHW				
NAT_NTOLINVPIND	% native non-tolerant invertivore individuals based on TOLERANCE_NRSA	FISHW				
NAT_NTOLINVPTAX	% native non-tolerant invertivore taxa based on TOLERANCE_NRSA	FISHW				
NAT_NTOLNTAX	No. native non-tolerant taxa based on TOLERANCE_NRSA	FISHW				
NAT_NTOLPIND	% native non-tolerant individuals based on TOLERANCE_NRSA	FISHW				
NAT_NTOLPTAX	% native non-tolerant taxa based on TOLERANCE_NRSA	FISHW				
NAT_OMNINTAX	No. native omnivore taxa	FISHW				
NAT_OMNIPIND	% native individuals as omnivore	FISHW	NONE			
NAT_OMNIPTAX	% native taxa as omnivore	FISHW	NONE			
NAT_PIND	native % of total individuals	FISHW		0	100	
NAT_PTAX	native % of total taxa	FISHW		0	100	
NAT_RBCATONTAX	No. native round-bodied sucker taxa	FISHW				
NAT_RBCATOPIND	% native individuals as round-bodied sucker	FISHW	NONE			
NAT_RBCATOPTAX	% native taxa as round-bodied sucker	FISHW	NONE			
NAT_RHEONTAX	No. native rheophilic taxa	FISHW				
NAT_RHEOPIND	% native individuals as rheophilic	FISHW	NONE			
NAT_RHEOPTAX	% native taxa as rheophilic	FISHW	NONE			
NAT_SALMNTAX	No. native salmonid taxa	FISHW				
NAT_SALMPIND	% native individuals as salmonid	FISHW	NONE			
NAT_SALMPTAX	% native taxa as salmonid	FISHW	NONE			
NAT_SENSLOTNTAX	No. native sensitive lotic  taxa	FISHW		0		
NAT_SENSLOTPIND	% native individuals as sensitive lotic	FISHW	NONE	0	100	
NAT_SENSLOTPTAX	% native taxa as sensitive lotic	FISHW	NONE	0	100	
NAT_SENSMIGRNTAX	No. native sensitive migratory taxa	FISHW		0		
NAT_SENSMIGRPIND	% native individuals as sensitive migratory	FISHW	NONE	0	100	
NAT_SENSMIGRPTAX	% native taxa as sensitive migratory	FISHW	NONE	0	100	
NAT_TOLRNTAX	No. distinct Tolerant native taxa based on TOLERANCE_NRSA	FISHW		0		
NAT_TOLRPIND	% Tolerant native individuals based on TOLERANCE_NRSA	FISHW		0	100	
NAT_TOLRPTAX	% Tolerant native taxa based on TOLERANCE_NRSA	FISHW		0	100	
NAT_TOTLNIND	Total native individuals	FISHW		0		
NAT_TOTLNTAX	Total native taxa	FISHW		0		
NAT_WTD_TV	Weighted average tolerance value among natives	FISHW				
NTOLBENTNTAX	No. non-tolerant benthic taxa	FISHW				
NTOLBENTPIND	% non-tolerant benthic individuals	FISHW				
NTOLBENTPTAX	% non-tolerant benthic taxa	FISHW				
NTOLCARNNTAX	No. non-tolerant carnivore taxa	FISHW		0		
NTOLCARNPIND	% non-tolerant carnivore individuals	FISHW		0	100	
NTOLCARNPTAX	% non-tolerant carnivore taxa	FISHW		0	100	
NTOLINVNTAX	No. non-tolerant invertivore taxa	FISHW		0		
NTOLINVPIND	% non-tolerant invertivore individuals	FISHW		0	100	
NTOLINVPTAX	% non-tolerant invertivore taxa	FISHW		0	100	
NTOLNTAX	no. non-tolerant (TOLERANCE_NRSA=(S)ensitive/(I)ntermediate) taxa	FISHW	NONE	0		
NTOLPIND	% non-tolerant (TOLERANCE_NRSA=(S)ensitive/(I)ntermediate) individuals	FISHW	NONE	0	100	
NTOLPTAX	% non-tolerant (TOLERANCE_NRSA=(S)ensitive/(I)ntermediate) taxa	FISHW	NONE	0	100	
OMNINTAX	No. omnivore taxa	FISHW		0		
OMNIPIND	% omnivore individuals	FISHW		0	100	
OMNIPTAX	% omnivore taxa	FISHW		0	100	
RBCATONTAX	No. round-bodied sucker taxa	FISHW				
RBCATOPIND	% round-bodied sucker individuals	FISHW				
RBCATOPTAX	% round-bodied sucker taxa	FISHW				
RHEONTAX	No. rheophilic taxa	FISHW				
RHEOPIND	% rheophilic individuals	FISHW				
RHEOPTAX	% rheophilic taxa	FISHW				
RT_NRSA	Reference/So-so/Trashed designations for sites used in NRSA					
SALMNTAX	No. Salmonid taxa	FISHW		0		
SALMPIND	% Salmonid individuals	FISHW		0	100	
SALMPTAX	% Salmonid taxa	FISHW		0	100	
SAMPLE_TYPE	Sample type					
SENSINVNTAX	No. sensitive invertivore taxa	FISHW		0		
SENSINVPIND	% sensitive invertivore individuals	FISHW		0	100	
SENSINVPTAX	% sensitive invertivore taxa	FISHW		0	100	
SENSNTAX	No. sensitive (TV<2.4) taxa	FISHW		0		
SENSPIND	% sensitive (TV<2.4) individuals	FISHW		0	100	
SENSPTAX	% sensitive (TV<2.4) taxa	FISHW		0	100	
SENSRHEONTAX	No. sensitive rheophilic taxa	FISHW		0		
SENSRHEOPIND	% sensitive rheophilic individuals	FISHW		0	100	
SENSRHEOPTAX	% sensitive rheophilic taxa	FISHW		0	100	
SITE_CLASS	"Site class (NAWQA, PROB, HAND)"					
SITE_ID	Site identification code					
STRAHLERORDER	Strahler stream order from RF3 stream data					
TOLRNTAX	No. distinct Tolerant taxa based on TOLERANCE_NRSA	FISHW		0		
TOLRPIND	% Tolerant individuals based on TOLERANCE_NRSA	FISHW		0	100	
TOLRPTAX	% Tolerant taxa based on TOLERANCE_NRSA	FISHW		0	100	
TOTLNIND	Total no. individuals	FISHW		0		
TOTLNTAX	Total no. distinct taxa	FISHW		0		
UID	Unique site visit ID					
VISIT_NO	Visit number for that year					
WTD_TV	"Hilsenhoff Biotic Index (HBI), sum of TV_i*prop_i, where TV_i=tolerance value for species i and prop_i=proportion of taxon i in sample, based on all taxa in sample"	FISHW	NONE	0	10	
YEAR	Sampling year					
